
A trochophore (/ ˈtroʊkəˌfɔːr, ˈtrɒ -, - koʊ -/; also spelled trocophore) is a type of free-swimming planktonic marine larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
Does trochophore have cilia?
The classic trochophore – a larval form shared with some other lophotrochozoans (e.g., molluscs) – is a top-shaped free-swimmer that moves through the water propelled by characteristic bands of cilia and usually has a sensory apical tuft of cilia (Fig. 2).
What is the function of trochophore?
trochophore, also called trochosphere, small, translucent, free-swimming larva characteristic of marine annelids and most groups of mollusks. Trochophores are spherical or pear-shaped and are girdled by a ring of cilia (minute hairlike structures), the prototroch, that enables them to swim.
What is the significance of trochophore larva?
Trochophore larva has a prominent status in the evolutionary context. Studies are suggestive of the Trochophore larva as a bridge between radial and bilateral symmetry. The larval form shows similarities with many invertebrate groups.
What is the definition of a trochophore?
Definition of trochophore : a free-swimming ciliate larva occurring in several invertebrate groups (such as the polychaete worms and mollusks)
How do you pronounce trochophore?
0:051:00How To Say Trochophore - YouTubeYouTubeStart of suggested clipEnd of suggested clipYojoa yojoa sojo fue yo voy sofofa yojoa.MoreYojoa yojoa sojo fue yo voy sofofa yojoa.
What animal phylum has trochophore larva?
Trochophores exist as a larval form within the trochozoan clade, which include the entoprocts, molluscs, annelids, echiurans, sipunculans and nemerteans. Together, these phyla make up part of the Lophotrochozoa; it is possible that trochophore larvae were present in the life cycle of the group's common ancestor.
Do flatworms have trochophore larvae?
The flatworms possess neither a lophophore nor trochophore larvae, although the larvae of one group of flatworms, the Polycladida (named after its many-branched digestive tract), are considered to be homologous to trochophore larvae.
Do arthropods have trochophore larvae?
Until very recently, the Arthropoda (insects & crustaceans) were considered possible close relatives of the Annelida, based on the fact that both groups are segmented, but no arthropod has a trochophore larva and no molecular studies support a close relationship.
What is the larval stage of arthropods?
This larval stage, called a nauplius, is found in barnacles, copepods, penaeid shrimps, and ostracods. It swims by using its jointed appendages that are moved by muscle contraction. These larvae have a nauplier eye, made up of photoreceptor cells that are thought to give rise to the photoreceptors in some adult forms.
What is called Swarm?
1 : a large number of bees that leave a hive together to form a new colony elsewhere. 2 : a large number grouped together and usually in motion a swarm of mosquitoes a swarm of tourists. swarm. verb. swarmed; swarming.
What is the difference between trochophore and veliger?
Unlike the trochophore, the newly hatched veliger may have or will develop many of the characteristic features of the adult including such structures as a muscular foot, eyes, rhinophores, a fully developed mouth, and a spiral shell (in fact, the veliger of nudibranchs has a shell, although the adult does not).
What is Prostomium in zoology?
: the portion of the head of an annelid worm (such as an earthworm) that is situated in front of the mouth.
Which of the following is the function of tubes in annelids?
Tubes are coiled to increase surface area for secretion or absorption in annelids.
What is the larva of Mollusca?
veliger, larva typical of certain mollusks such as marine snails and bivalves and a few freshwater bivalves. The veliger develops from the trochophore (q.v.) larva and has large, ciliated lobes (velum).
What is the larva of Arthropoda?
This larval stage, called a nauplius, is found in barnacles, copepods, penaeid shrimps, and ostracods. It swims by using its jointed appendages that are moved by muscle contraction. These larvae have a nauplier eye, made up of photoreceptor cells that are thought to give rise to the photoreceptors in some adult forms.
What are the basic characteristics of molluscs?
Mollusca CharacteristicsThey are mostly found in marine and freshwater. ... They exhibit organ system level of organization.Their body has a cavity.The body is divided into head, visceral mass, muscular foot and mantle.The head comprises of tentacles and compound eyes.The body is covered by a calcareous shell.More items...•
What is the function of cilia in the human body?
Almost every cell in the human body contains cilia. These cells are responsible for moving molecules and lipids past the cell. This process helps keep the cells and human body functioning properly. Cilia are also found on eukaryotic cells and help eukaryotic cells move. Many unicellular Protozoa have cilia on the cell.
What is the structure of the cilia?
Cilia also have a unique molecular structure. Inside the cilia, a microtubular cytoskeleton backbone is surrounded by a plasma membrane called the ciliary axoneme. Each ciliary axoneme has nine microtubular pairs; however, the arrangement of the microtubular pairs is different in motile cilium and non-motile cilium. In motile cilium, there is a microtubular pair found in the center (9 + 2), but in the non-motile cilium, there are no centrally located microtubular pairs (9 + 0).
What is Cilia?
Cilia are small appendages that whip back and forth in eukaryotic cells. The primary purpose of cilia is to help a cell move in cellular fluid and help particles move past the cell in one direction, accomplishing this by their back and forth movements. However, cilia are only about 0.1 millimeters in size. In the human body, cilia are found in almost all of the cells.
What is the plural of cilia?
In Latin, the term cilia means eyelashes and is the plural form of a cilium. Thus, a group of tiny hair-like structures on a cell is called cilia, whereas one or two hair-like structures are called flagellum. There are also synonyms for cilia and cilium, undulipodium being the synonym for cilium and undulipodia being the synonym for cilia. Therefore, undulipodia is a group of tiny appendages that can be found on eukaryotic cells.
What is the term for when the cilia work together to help move particles past the cell?
Intraflagellar transport is when the cilia work together to help move particles past the cell. When the cilia are not functioning correctly, it can affect the rest of the cell. Ciliopathy is when the cilia are not working correctly.
What are the two main structures of the cilia?
Tubulin and basal bodies make up the two main structures of cilia. Tubulin are the hair-like structure of the cilia and are composed of small protein pieces connected to the cell by basal bodies. These protein pieces are made in the cell and delivered to the tubulin.
Where are the cilia found?
The motile cilia are commonly found in unicellular organisms and the respiratory tract, while non-motile cilia are found in mammalian cells and are used for sensory purposes. The nodal cilia are located in the early stages of embryo development.
What is the ring of cilia that enables a troch to swim?
Trochophores are spherical or pear-shaped and are girdled by a ring of cilia (minute hairlike structures), the prototroch, that enables them to swim.
What is the second stage of a trochophore?
In some mollusks (such as gastropods and bivalves), the trochophore develops into a second stage, the veliger ( q.v. ), before metamorphosing to adult form. Rotifers and the larvae (sometimes considered trochophores) of such invertebrates as phoronids and bryozoans are trochophore-like in appearance.
What are the structures that are below the prototroch?
Below the prototroch are the mouth, stomach, anus, and other structures including the solenocyte, the function of which seems to be to maintain proper internal salt-water balance, and, in some species, one or two additional ciliary rings.
What is the control of the pre-oral cilia?
The control of beating of the pre-oral cilia is probably due to a mechanism similar to that in gastropod veligers investigated by a number of authors reviewed by Arkett et al. (1987). They showed, in veligers of the gastropod Calliostoma, that the cells bearing the pre-oral cirri are connected by gap junctions and are thus electrically coupled. Propagated Ca 2+ -dependent action potentials are initiated from the cerebral ganglion, causing velum-wide arrests of ciliary beating, while reduction in beating rate is accompanied by a slow depolarisation of the cell membrane. Direct evidence of excitatory nervous control by sensory serotinergic cells in the probable homologue of the apical sense organ of marine gastropods has been obtained from laser ablation studies with pulmonate embryos ( Kuang and Goldberg, 2001 ). The velum of Mytilus larvae has dual innervation: excitatory serotinergic and inhibitory dopaminergic ( Beiras and Widdows, 1995) and acetylcholinesterase occurs in the ciliated cells of the velum ( Raineri, 1995 ). Immunofluorescence studies suggest nervous control of the ciliated cells by catecholaminergic cells on the rim of the velum, but no rim to apical organ connections has yet been located ( Croll et al., 1997 ). The abrupt cessation of beating and changes in beating rate that occur in Pecten maximus veligers ( Cragg, 1989) suggest nervous control similar to that revealed by Beiras and Widdows (1995).
What are the reproductive characteristics of molluscs?
Molluscs have extreme variation in reproductive characteristics. A generic mollusc is dioecious with paired gonads. Eggs or sperm are released into the coelomic cavity and carried outside by ducts. Fertilization is external and the zygote develops into a gastrula and then a free-swimming trochophore larva. Trochophore larvae develop into a veliger in marine taxa. The veliger larvae have a well-developed foot, shell, and other structures. A veliger will settle on the bottom of the water body and metamorphose into the adult form. In freshwater unionid bivalves, larvae develop into glochidia that are brooded and later released to attach to fish hosts used for dispersal; others have direct development into adults (Sphaeriidae or pill clams); while some retain the veliger larval form (e.g., Dreissena or zebra mussels). Freshwater gastropods also have great variation in developmental patterns. Gastropods can be oviparous, ovoviviparous, or viviparous. Larval forms are present in some taxa. For example, the neritids are oviparous and deposit eggs on substrates that hatch into a swimming larval form. The larvae undergo metamorphosis into the adult form. Oviparous taxa have direct development, while ovoviviparous species brood their young and release them at a larger size.
How long do sipunculans stay in plankton?
The sipunculans follow the first trochophore with a prolonged larval stage in which they may stay in the plankton for several months. If an early trochophore happens to hit the substrate prematurely, it will not metamorphose, but will most likely die.
What are the characteristics of a mollusk?
Molluscs are protostomous coelomates, exhibiting spiral, determinate cleavage and schizocoely as well as having trochophore larvae and the blastopore forming the mouth of the adult. They usually possess all or a combination of the following features: (1) a reduced coelom and vestiges of metamerism; (2) a mantle or fleshy epidermis of the dorsal body wall which has glands capable of secreting calcium carbonate to form an exoskeletal shell or shelly parts, such as plates, spines, and spicules; (3) a mantle cavity or an invagination of the mantle which contains a pair or more of specialized respiratory structures, the ctenidia or gills, and into which the digestive, metanephridial excretory, and reproductive systems debouch their products; (4) the ventral body surface modified into a pedal groove or muscularized foot for progression or locomotion; (5) a special chitinized, rasp-like tongue or radula; (6) an open hemocelic circulatory system with a chambered heart having auricles and ventricles. Additionally, the nervous system has variously paired ganglionic portions, particularly cerebral, pedal, and visceral ones, as well as ventral anteroposterior cord-like connective elements; specialized sensory structures were evolved for olfaction, vision, balance, and tactile stimulation.
Is the apical organ a sense organ?
The structure of the apical organ suggest s that it is a sense organ, but a sensory role has not been demonstrated experimentally, though in the trochophore and early veliger the nervous system has yet to differentiate. In later veligers, broad conclusions about function can be drawn on the basis of structural similarities with more thoroughly investigated organisms. The complex brush border of the apical organ, with vertical and horizontal microvilli interwoven with extracellular material, is similar to that of molluscan chemosensory epithelia on opisthobranch rhinophores (Goebbeler and Klussmann-Kolb, 2007 ). The cells bearing the cilia forming the apical tuft resemble sensory neurons in such epithelia. The ciliated cavity cells of the apical organ resemble cells in a range of putative or known chemosensory structures: the apical sensory organ of gastropod veligers ( Kempf and Page, 2005; Page and Kempf, 2009 ), the rhinophore of Nautilus ( Ruth et al., 2002 ), and the ampullary system of chitons ( Haszprunar et al., 2002 ). The sensory modality of this cell type remains to be established in many cases, though Lucero et al. (1992) have detected an electrical response to chemical stimuli from similar cells in squid. Detailed studies with the homologous apical sensory organ of gastropod larvae are also illuminating. Ablation of the cells with ciliated cavities in the apical sensory organ of a nudibranch veliger prevents metamorphosis in response to settlement cues ( Hadfield et al., 2000 ). Receptors for such cues have been located on cilia harvested from metamorphically competent veligers of a prosobranch ( Baxter and Morse, 1992 ).
Who discovered the trochophore?
Historical Retrospect of Trochophore Larva: 1. Loven (1840), a Swedish naturalist, was the first man who discovered trochophore larva. Since then the larva was known as Loven’s larva. 2. Semper (1859) used the name Trochosphaera, a rotifer for the organism. ADVERTISEMENTS:
Who gave the name trochophore?
Ray Lankester (1877) gave the name ‘Trochophore’ to this larval form. 4. Afterwards it was Hatschek (1879) who also supported the most accepted name ‘Trochophore’. 5. Hyman (1957) and Barnes (1980) tried to establish relationship of trochophore with other groups of animals.
What is the ciliary band in the body called?
7. A circlet of strong locomotor ciliary band, called the pre-oral circlet or Prototroch (Fig. 17.12A), encircle the body around the middle and lies above the mouth, and the metatroch which is a ciliary transverse ring behind the mouth.
Which trochrome exhibits no metamerism?
11. Trochophore exhibits no metamerism and the rudiment of future adult trunk is seen as a small region at the posterior pole.
What is the name of the tuft of cilia?
16. At the apical end, a number of long cilia emerge from the apical plate, called apical tuft of cilia.
Do trochophore larvae eat eggs?
In some other groups (e.g., Sipunculans, nereids and eunicids of polychaetes), the trochophore larvae are lecithotrophic larvae. They do not take any food from external sources and mainly feed upon the yolk originally laid down in the egg. The lecithotrophic larvae lead a short planktonic life. Metamorphosis of the larvae is best seen in Polygordius.
Where does prototroch come from?
8. The prototroch arises from special cells, called trochoblasts.
What are the functions of the cilia?
Cilia Function. The important functions performed by cilia involve locomotion and sensory functions. They play a major role in cell cycle and replication and also in the development of humans and animals. Multiple cilia move in a rhythmic motion that keeps the internal passageways free from mucus or any foreign agent.
What are Cilia?
Cilia are small, slender, hair-like structures present on the surface of all mammalian cells. They are primitive in nature and could be single or many.
What are the two types of cilia?
The organelle cilia are found in eukaryotic cells. They can be of two types – motile cilia and non-motile cilia. The non-motile cilia are known as primary cilia and act as sensory organelles. Cilia are structurally identical to flagella.
What are primary cilia?
Primary Cilia are non-motile cilia that were first discovered in 1898. These structures were long believed to be vestigial organelles. However, recent researches presented the biological roles of primary cilia that they function as a sensory cellular antenna that coordinates a large number of cellular signalling pathways. Apart from these, they also help in: 1 Proper urine flow by signalling the kidney cells. 2 They act as mechanoreceptors or sensory receptors. 3 The cilia function by permitting the transfer of important particles from one side of the light-sensitive cells to another in the retina.
How big is the cilia?
They are very minute structures ranging from 0.25μm in diameter to 20μm in length.
What is the name of the disorder in which the cilia do not function normally?
Dysfunction or defects in primary and motile cilia are known to cause numerous distressing genetic disorders known as ciliopathies. Primary Ciliary Dyskinesia: It is an autosomal recessive disorder in which the cilia do not function normally.
What is a non-motile cilia?
Non-motile Cilia. Primary Cilia are non-motile cilia that were first discovered in 1898. These structures were long believed to be vestigial organelles. However, recent researches presented the biological roles of primary cilia that they function as a sensory cellular antenna that coordinates a large number of cellular signalling pathways.
Why do trochophores move?
The cilia on a trochophore are moved rapidly in order to create a water eddy, thus controlling the direction of their movement.
What is a trochophore?
A trochophore is a free-swimming, ciliated larva of many worms and some mollusks.
What is the fluid in the circulatory system of spiders?
Hemolymph is a fluid in the circulatory system of some arthropods (including spiders, crustaceans such as crabs and shrimp, and even some insects such as stoneflies). The hemocoel is a cavity in which the hemolymph flows.
What is the function of the earthworm's segments?
Segmentation helps the earthworm move, where each segment has muscles and bristles called setae . The bristles or setae help anchor and control the worm when moving through soil. The bristles hold a section of the worm firmly into the ground while the other part of the body protrudes forward.
What is a tonguelike organ that is covered with chitinous teeth and that is used for feeding by?
a rasping, tonguelike organ that is covered with chitinous teeth and that is used for feeding by many mollusks
What is a mollusk that has tentacles extending from the head?
a marine mollusk that has tentacles extending from the head; an octopus, squid, cuttle-fish, or nautilus
Do protostomes have a coelom?
Both are protostomes and have a true coelom ( body cavity). Both have a similar larval stage, bilateral symmetry, and organ systems

Overview
A trochophore is a type of free-swimming planktonic marine larva with several bands of cilia.
By moving their cilia rapidly, they make a water eddy, to control their movement, and to bring their food closer, to capture it more easily.
Occurrence
Trochophores exist as a larval form within the trochozoan clade, which include the entoprocts, molluscs, annelids, echiurans, sipunculans and nemerteans. Together, these phyla make up part of the Lophotrochozoa; it is possible that trochophore larvae were present in the life cycle of the group's common ancestor.
Etymology
The term trochophore derives from the ancient Greek τροχός (trókhos), meaning "wheel", and φέρω (phérō) — or φορέω (phoréō) —, meaning 'to bear, to carry', because the larva is bearing a wheel-shaped band of cilia.
Feeding habits
Trochophore larvae are often planktotrophic; that is, they feed on other plankton species.
Life cycle
The example of the development of the annelid Pomatoceros lamarckii (family Serpulidae) shows various trochophore stages (image: D-F): D - early trochophore ; E - complete trochophore ; F - late trochophore ; G - metatrochophore.
External links
• Nielsen, Claus (2018-07-01). "Origin of the trochophora larva". Royal Society Open Science. 5 (7): 180042. Bibcode:2018RSOS....580042N. doi:10.1098/rsos.180042. ISSN 2054-5703. PMC 6083724. PMID 30109065.