Knowledge Builders

where are bryozoans found

by Amanda Feest Published 3 years ago Updated 2 years ago
image

Worldwide, bryozoans are found on every continent except Antarctica. Although most bryozoans are marine, one class (Phylactolaemata) lives only in freshwater. About 20 freshwater species occur on our continent. These usually prefer the rather quiet waters of lakes, ponds, and swamps, but some live in streams.

Full Answer

Do bryozoans live in lakes?

Worldwide, bryozoans are found on every continent except Antarctica. Although most bryozoans are marine, one class (Phylactolaemata) lives only in freshwater. About 20 freshwater species occur on our continent. These usually prefer the rather quiet waters of lakes, ponds, and swamps, but some live in streams.

How many types of bryozoan are there?

There are over 4,000 species found worldwide, with about 50 species living solely in freshwater. Physical description: Each species of bryozoan can have a different appearance, but perhaps the one that stands out the most is Pectinatella magnifica. Their large gelatinous colonies are eye catching and intriguing.

Where can I find media related to Bryozoa?

The Biology of Bryozoans, Academic Press, New York. Wikimedia Commons has media related to Bryozoa.

Where are bryozoans found in Kansas?

In Kansas, fossil bryozoans are common in the Pennsylvanian and Permian rocks of the eastern part of the state. Bryozoan fossils from the Topeka Limestone in Kansas.

image

How are bryozoans formed?

Bryozoan colonies start out with a single zooid, which may be produced either sexually or asexually (through budding) by the parent colony. As this original zooid begins feeding, it buds to form additional genetically identical zooids. These new zooids also bud, forming the colony.

Where are bryozoan fossils found?

Bryozoan fossils can be found in Kentucky's Ordovician, Silurian, Devonian, Mississippian, and Pennsylvanian rocks. Fenestrate bryozoa colonies, like the three diffenent types shown above, are lace-like in construction. The individual bryozoan animals lived in microscopic tubes or pores on the lace branches.

What period did bryozoans live in?

Ordovician periodThe oldest fossils of bryozoans, colonies made of tiny individual animals called zooids, were previously dated to the Ordovician period around 480 million years ago. This is about 50 million years later than most other animal groups first emerged.

Are bryozoans animals or plants?

Moss animals, or bryozoans, are actually animals in a plant disguise! They can be found attached to hard surfaces like rocks or pilings — you will often see tufts of them on floating docks or small white patches of them encrusting kelp fronds.

When did bryozoans go extinct?

Fossil range These bryozoans lived from the Carboniferous period (Tournaisian age) to the Permian period (Leonard age) (345.3 to 268.0 Ma), when this genus became extinct.

How did bryozoans go extinct?

Stenolaemates—and bryozoans, in general—were hard hit by the end-Permian extinction and are rare in Triassic-aged rocks. All orders of Stenolaemata, except Cyclostomata, went extinct by the end of the Triassic.

Are bryozoans edible?

A bryozoan colony, consisting of individuals called zooids, may resemble a brain-like gelatinous mass and be as big as a football, and can usually be found in shallow, protected areas of lakes, ponds, streams and rivers, and is often attached to things like a mooring line, a stick, or a dock post, etc.” While Bryozoans ...

Should I remove bryozoan?

Bryozoans are a food source to fish, insects and snails. They also clean the water as they consume microorganisms, which typically results in good water quality. Due to their positive impact on ponds, removal is unnecessary.

Are bryozoans important?

Both living and fossil bryozoans can be found in the British Isles. Bryozoans are important because they are: Foulers. Bryozoans can affect the performance or function of human-made structures such as oil rigs, buoys, moorings, ship hulls and intake pipes for power stations.

What is this jelly like blob under my dock?

Typically, when people ask this question, what they have discovered is a colony of the freshwater bryozoan Pectinatella magnifica. Freshwater bryozoans are microscopic aquatic invertebrates that live in colonies that can form into jelly-like clumps, and are often found attached to docks or sticks.

Are bryozoans harmful?

Montz says bryozoans are quite common in many Minnesota waters, ranging from large rivers to lakes to small ponds. They are not toxic, venomous, or harmful. They don't really seem to cause problems for people, except for the "ick" factor and occasionally clogging underwater screens or pipes.

What are bryozoans eaten by?

Bryozoans eat microscopic organisms and are eaten by several larger aquatic predators, including fish and insects. Snails graze on them, too. Like mussels and other filter feeders, bryozoans gradually cleanse the water as they feed.

Where are horn coral fossils found?

Horn corals are an extinct order of coral known as Rugosa. Abundant horn coral fossils can be found in the Confusion Range in Millard County. During the Triassic Period (~215 mya), central Utah was a transition zone between river flood plains to the southeast and seas to the northwest.

What type of rock is bryozoan limestone?

Bryozoan limestone is a formation of Danian limestone, generally found underlying the younger Copenhagen formation and above the Cretaceous chalk. The mineralogy between the Danian and Cretaceous formations is similar, resulting in similar mechanical responses of the matrix material.

What is a bryozoan blob?

Bryozoans are aquatic invertebrates like jellyfish, but unlike jellies, which are singular organisms, this blob is made up of thousands of individual microscopic animals, called zooids, living in a colony.

What is Graptolite fossil?

Fossil graptolites are thin, often shiny, markings on rock surfaces that look like pencil marks, and their name comes from the Greek for 'writing in the rocks'.

How many species of bryozoan are there?

Their name, bryozoan, translates from Latin as “moss animal”. There are over 4,000 species found worldwide, with about 50 species living solely in freshwater.

What does it mean when you have bryozoans in your pond?

If you have bryozoans in your pond, it is an indicator that you have a healthy ecosystem. Bryozoan feeding habits mean they filter the water as they feed, like oysters in saltwater, helping to consume algae and remove suspended sediments.

What are the colonies of P. magnifica?

Their large gelatinous colonies are eye catching and intriguing. What we see in the water is the colony made up of hundreds to thousands of microscopic animals, called zooids. In P. magnifica, the zooids grow in a unique rosette pattern on a substrate they create themselves; each rosette contains several zooids.

When do statoblasts disintegrate?

They exhibit rapid growth with warm weather and are sensitive to drops in the water temperature1, 3. Colonies typically begin to disintegrate in early September, as the season begins to change1. When a colony disintegrates, the statoblasts are released and moved to a new location by water currents or on another organism, or they remain in the sediment until the following spring.3

Where is P. magnifica found?

Geographic range: The native range of P. magnifica is east of the Mississippi River, as far north as Ontario, and as far south as Florida. Its range has expanded westward into Texas and the Pacific Northwest, possibly due to effects from climate change and movement of ships.1, 3 It has been found in Japan and Korea, likely as a result of waterfowl migrations.3 P. magnifica seems to prefer nutrient rich (eutrophic) waters with low turbidity3, although some studies have shown that it prefers lower nutrient waters.1.

Can bryozoans clog ponds?

During a particularly “good year” they can form a bloom, and like algae, too much can be a bad thing. In the case of bryozoans, during a large bloom they can clog inflow and outflow pipes and water filters. There is also some research to show they can harbor a parasite ( Tetracapsuloides bryosalmonae) that can impact salmonids, especially Rainbow Trout2. They are not a threat to fisheries in South Carolina.

Who wrote the book Ecology and Classification of North American Freshwater Invertebrates?

Ecology and Classification of North American Freshwater Invertebrates, by James H. Thorp and Alan P. Covich, Academic Press, 2001, pp. 505–526. If this document didn’t answer your questions, please contact HGIC at [email protected] or 1-888-656-9988.

What is the name of the bryozoan?

Common name: magnificent bryozoan. Taxonomy: available through. www.itis.gov. Identification: Pectinatella magnifica is a species of freshwater bryozoan in the class Phylactolaemata. Like other species of bryozoans (also known as Ectoprocta or commonly as moss animals), the individual microscopic aquatic invertebrates (called a zooid) ...

What are the tentacles of a P. magnifica zooid?

The individual P. magnifica zooids have mucous-coated tentacles that trap diatoms, phytoplankton, and other microscopic organisms, where cilia, or tiny hairs lining the tentacles, sweep the food to the mouth (Ricciardi and Reiswig 1994, Wood 2010). Individual zooids may filter an average of 8.8 ml of water/ day (Bullivant 1968).

What parasites are associated with Pectinatella magnifica?

Pectinatella magnifica can serve as an intermediate host of Tetracapsuloides bryosalmonae, a myxozoan parasite that causes Proliferative Kidney Disease (PKD) in salmonids (Okamura and Wood 2002). PKD is especially problematic for rainbow trout ( Oncorhynchus mykiss) aquaculture in the western U.S. (Merck 2015). References.

Where is Pectinatella native to?

Native Range: Pectinatella magnifica is native to freshwater drainages of eastern North America from New Brunswick and Ontario, Canada; south to Florida and Mississippi. The native distribution spans throughout the Great Lakes region east of the Mississippi River mainstem (Pennak 1989, Wood 2010). Table 1.

Where can I find Pectinatella magnifica?

Ecology: Pectinatella magnifica are typically found in lentic environments including freshwater lakes, ponds, and swamps, but can also be found in lotic environments (i.e., streams). The colonies are sessile with little preference among attaching surfaces (Joo et al. 1992), frequently found attached to rocks, wood, aquatic vegetation, pilings, or other submerged surfaces. On rare occasions the colonies will be found free-floating (Ricciardi and Reiswig 1994, Wood 2010, Joo et al. 1992). Pectinatella magnifica has been defined as a warm water species (Ricciardi and Reiswig 1994) although the species can display a wide temperature tolerance (4-32 °C) (Everitt 1975, Ricciardi and Reiswig 1994), but cannot tolerate saline waters (Everitt 1975).

What is the name of the new group of bryozoa?

The new group was given the name " Entoprocta ", while the original Bryozoa were called "Ectoprocta". Disagreements about terminology persisted well into the 20th Century, but "Bryozoa" is now the generally accepted term. Colonies take a variety of forms, including fans, bushes and sheets.

What is the feeding structure of a bryozoan?

Typically about 0.5 millimetres ( 1⁄64 inch) long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters.

How long are phoronids?

Phoronids resemble bryozoan zooids but are 2 to 20 cm (1 to 8 in) long and, although they often grow in clumps, do not form colonies consisting of clones. Brachiopods, generally thought to be closely related to bryozoans and phoronids, are distinguished by having shells rather like those of bivalves.

What is the relationship between a lophophorate and a mollusc?

Molecular phylogeny analyses from 1995 onwards, using a variety of biochemical evidence and analytical techniques, placed the lophophorates as protostomes and closely related to annelids and molluscs in a super-phylum called Lophotrochozoa. "Total evidence" analyses, which used both morphological features and a relatively small set of genes, came to various conclusions, mostly favoring a close relationship between lophophorates and Lophotrochozoa. A study in 2008, using a larger set of genes, concluded that the lophophorates were closer to the Lophotrochozoa than to deuterostomes, but also that the lophophorates were not monophyletic. Instead, it concluded that brachiopods and phoronids formed a monophyletic group, but bryozoans (ectoprocts) were closest to entoprocts, supporting the original definition of "Bryozoa".

What is the common name for moss animals?

Works since 2000 have used various names to resolve the ambiguity, including: "Bryozoa", "Ectoprocta", "Bryozoa (Ectoprocta)", and "Ectoprocta (Bryozoa)". Some have used more than one approach in the same work. The common name "moss animals" is the literal meaning of "Bryozoa", from Greek βρυόν ('moss') and ζῷα ('animals').

What is the phylum of moss?

Ectoprocta (Nitsche, 1869) (formerly subphylum of Bryozoa) Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about 0.5 millimetres ( 1⁄64 inch) long, they have a special feeding structure called a lophophore, ...

How many statoblasts does Plumatella repens produce?

A study estimated that one group of colonies in a patch measuring 1 square meter (11 square feet) produced 800,000 statoblasts. Cupuladriid Bryozoa are capable of both sexual and asexual reproduction.

Where are bryozoan fossils from?

Bryozoan fossils from the Topeka Limestone in Kansas. These fossils were deposted during the Pennsylvanian Subperiod, about 300 million years ago, and illustrate the branching and netlike forms of some bryozoan colonies. Freshwater bryozoans are virtually unknown as fossils, presumably because they did not have mineralized skeletons.

Where are marine bryozoans found?

Marine bryozoans show up in the fossil record in the early part of the Ordovician Period, about 485 million years ago. In Kansas, fossil bryozoans are common in the Pennsylvanian and Permian rocks of the eastern part of the state. Bryozoan fossils from the Topeka Limestone in Kansas. These fossils were deposted during the Pennsylvanian Subperiod, ...

What are the members of a bryozoan colony called?

Individual members of a bryozoan colony are called zooids. Although all zooids in the colony are physically connected, each lives in its own calcium carbonate compartment. Some or all zooids in a colony are feeding zooids, equipped with tentacle-bearing feeding organs. Bryozoans feed by projecting these tentacles into the water through openings in their external skeletons. The tentacles have tiny moving filaments called cilia, which create currents that draw microscopic organisms and plants into the mouth. Feeding zooids also have an alimentary canal, muscles, a nervous system and a U-shaped digestive tract. Simple egg- and sperm-producing structures are also present in some zooids in every colony.

How do bryozoan colonies start?

Bryozoan colonies start out with a single zooid, which may be produced either sexually or asexually (through budding) by the parent colony. As this original zooid begins feeding, it buds to form additional genetically identical zooids. These new zooids also bud, forming the colony. Large colonies may consist of hundreds of thousands or even millions of zooids.

What kingdom are bryozoans in?

Taxonomic Classification: Bryozoans belong to Kingdom Animalia, Phylum Bryozoa. The phylum is divided into three classes, the Phylactolaemata (freshwater bryozoans), the Stenolaemata, and Gymnolaemata.

What are bryozoans made of?

Bryozoans are sometimes confused with corals, another colonial group of animals. Like corals, most bryozoans secrete external skeletons made of calcium carbonate, which form the framework of the colony. Bryozoans, however, are more complex organisms than corals and generally don't build reefs.

Why are bryozoans considered fossils?

Freshwater bryozoans are virtually unknown as fossils, presumably because they did not have mineralized skeletons. Throughout their nearly 500-million-year history, marine bryozoans have been abundant and widely distributed geographically. They are the most abundant fossils in many limestones, calcareous shales, and mudstones.

Where did the bryozoan come from?

A variety of fossil bryozoans from the Ordovician of Estonia. Image by Mark A. Wilson (public domain). Fossils from this phylum of “moss animals”—the translation of bryozoan from Greek—have been found as far back as the Early Ordovician (~480 million years ago).

Which bryozoan is the most abundant?

Gymnolaemates are the most abundant and diverse bryozoans today, comprising at least 80% of known bryozoan species. This abundance has only come about relatively recently, with the evolution of Cheilostomata in the Jurassic and diversification of this now-dominant order during the second half of the Cretaceous.

What was the dominant class of bryozoans during the Paleozoic?

During the Paleozoic, the Stenolaemata were the dominant class of bryozoans and an important component of early reef structures and benthic habitats. In addition to branching and encrusting forms (e.g., Polypora ), these early bryozoans also formed mound-like structures (e.g., Prasopora) and more eccentric forms such as the corkscrew-like Archimedes. Check out the 3D models below to explore these stenolaemates.

How many species of bryozoans are there?

To date, more than 17,800 species of fossil bryozoans have been described and more than 6,000 living species are known.

What are the predators that eat bryozoans?

In their various environments, bryozoans are occasionally consumed by predators, such as the nudibranch in the video below. Predators like nudibranchs, sea urchins, and sea stars move along the surface of a colony and consume individual zooids. Predation of this type can induce the formation of defensive structures (e.g., spines) by the colony. Anti-predatory defenses can be energetically costly to produce so some species (e.g., Membranipora membranacea) only produce them when there is an immediate threat (i.e., predator present). As demonstrated in laboratory experiments by Drew Harvell (1984), spines can take up to two days to form, meaning that individuals zooids will be consumed—before and while defenses are produced—but the colony will most likely survive. Predation of this type may have contributed to the evolution of specialized zooids like the vibricula and avicularia.

What are the different forms of zooid?

Zooids can take several forms (e.g., autozooids, avicularia, heterozooids, kenozooids, and vibracula) but the most common forms in each of the classes are autozooids, which function in feeding the colony and excreting waste. A generic autozooid is shown in the image below, on the right.

When did the first stenolaemata appear?

Stenolaemata are the first bryozoans to appear in the fossil record, dating to the Early Ordovician approximately 480 million years ago. Though they are the first in the record, they were not likely the first of the three classes to evolve. As discussed by Taylor and Waeschenbach (2015) in their review of the topic, the first Stenolaemata may have arisen through the biomineralization (i.e., skeletonization) of the outer wall of an early gymnolaemate. This is only one proposed hypothesis of relationship, however, as other phylogenies have placed Stenolaemata basal to the Gymnolaemata. Regardless of the relationship between these two classes, Phylactolaemata is very likely basal to both (see the phylogeny above). Within the Stenolaemata, a handful of orders were present during the Paleozoic, but only a single order, Cyclostomata, survived beyond the Triassic and to the present.

What are colonies of bryozoans called?

What folks are likely seeing are colonies of bryozoans, says DNR aquatic invertebrate biologist Gary Montz. These microscopic animals, sometimes referred to as moss animals, make jellylike tubes and attach themselves to sticks, rocks, and other submerged objects.

What happens to the bryozoan colony in the Baudette River?

Bryozoan colony attached to a rock in the Baudette River. Each fall the bryozoans begin to die off, but create overwintering "eggs" that will form new colonies the next year. When the colony is dying, gas produced by decomposition may cause it to float loose, sending gelatinous globs floating down the river.

Is a bryozoan poisonous?

Montz says bryozoans are quite common in many Minnesota waters, ranging from large rivers to lakes to small ponds. They are not toxic, venomous, or harmful. They don't really seem to cause problems for people, except for the "ick" factor and occasionally clogging underwater screens or pipes. "Just another of the really strange life forms ...

Where are bryozoans found?

Bryozoans are found in freshwater, brackish and marine ecosystems throughout the world, from all depths and latitudes. ( Brusca and Brusca, 2003; Ramel, 2012)

What is the habitat of a bryozoan?

Habitat. Bryozoans are sessile and colonial, typically settling on hard substrate including sand grains, rocks, and shells, as well as on blades of kelp or other algae, although some species settle on softer sediment.

What is the physical description of a zooid?

Physical Description. An individual organism within a colony is called a zooid, and is made up of a cystid and a polypide. The cystid is the outer casing (the chitinous, calcified or gelatinous zoecium, secreted by the zooid), and the attached body wall. The polypide is comprised of the lophophore and viscera.

How do bryozoan colonies grow?

A bryozoan colony begins with a single individual, known as an ancestrula. Ancestrulas are sexually produced, but colonies grow through asexual reproduction. Breeding is somewhat regulated by water temperatures and levels of sunlight: rising temperatures and increased light trigger phytoplankton growth which, in turn, triggers budding and, to a lesser extent, sexual reproduction. Species may free-spawn or, more often, females will brood eggs for at least a short time. Larvae of brooding species settle much more quickly following hatching, as their larval forms cannot feed. ( Brusca and Brusca, 2003; "Introduction to the Bryozoans", 2011)

What is the taxonomic classification of bryozoa?

The taxonomic classification of Bryozoa is complex and historically problematic. The name Bryozoa was first used by Christian Gottfried Ehrenberg in 1831, to refer to a group of organisms previously referred to as Polyzoa by J. Vaughan Thompson, just one year before.

What are the three classes of bryozoans?

Bryozoans have traditionally been placed into three classes: Phylactolaemata, Stenolaemata, and Gymnolaemata, which includes orders Ctenostomata and Cheilostomata. Though the majority of bryozoan species are marine, fresh and brackish water forms are also known.

How does a phytoplankton breed?

Breeding is somewhat regulated by water temperatures and levels of sunlight: rising temperatures and increased light trigger phytoplankton growth which, in turn, triggers budding and, to a lesser extent, sexual reproduction. Species may free-spawn or, more often, females will brood eggs for at least a short time.

image

Overview

Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about 0.5 millimetres (1⁄64 in) long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic …

Description

Bryozoans, phoronids and brachiopods strain food out of the water by means of a lophophore, a "crown" of hollow tentacles. Bryozoans form colonies consisting of clones called zooids that are typically about 0.5 mm (1⁄64 in) long. Phoronids resemble bryozoan zooids but are 2 to 20 cm (1 to 8 in) long and, although they often grow in clumps, do not form colonies consisting of clones. Brachiopods, …

Taxonomy

The phylum was originally called "Polyzoa", but this name was eventually replaced by Ehrenberg's term "Bryozoa". The name "Bryozoa" was originally applied only to the animals also known as Ectoprocta (lit. 'outside-anus'), in which the anus lies outside the "crown" of tentacles. After the discovery of the Entoprocta (lit. 'inside-anus'), in which the anus lies within a "crown" of tentacle…

Physiology

Most species are filter feeders that sieve small particles, mainly phytoplankton (microscopic floating plants), out of the water. The freshwater species Plumatella emarginata feeds on diatoms, green algae, cyanobacteria, non-photosynthetic bacteria, dinoflagellates, rotifers, protozoa, small nematodes, and microscopic crustaceans. While the currents that bryozoans generate to draw f…

Ecology

Most marine species live in tropical waters at depths less than 100 meters (330 ft; 55 fathoms). However, a few have been found in deep-sea trenches, especially around cold seeps, and others near the poles.
The great majority of bryozoans are sessile. Typically, sessile bryozoans live on hard substrates including rocks, sand or shells. Encrusting forms are much th…

See also

• International Bryozoology Association
• List of prehistoric bryozoan genera
• Colony (biology)

Bibliography

• Taylor, Paul D. (2020). Bryozoan Paleobiology. London, UK: Natural History Museum. ISBN 9781118455005.
• Hall, S.R.; Taylor, P.D.; Davis, SA; Mann, S (2002). "Electron diffraction studies of the calcareous skeletons of bryozoans". Journal of Inorganic Biochemistry. 88 (3–4): 410–419. doi:10.1016/S0162-0134(01)00359-2. PMID 11897358.

External links

• Index to Bryozoa Bryozoa Home Page, was at RMIT; now bryozoa.net
• Other Bryozoan WWW Resources
• International Bryozoology Association official website
• Neogene Bryozoa of Britain

1.Bryozoans (Moss Animals) | Missouri Department of …

Url:https://mdc.mo.gov/discover-nature/field-guide/bryozoans-moss-animals

1 hours ago Worldwide, bryozoans are found on every continent except Antarctica. Although most bryozoans are marine, one class (Phylactolaemata) lives only in freshwater. About 20 freshwater species occur on our continent. These usually prefer the rather quiet waters of lakes, ponds, and …

2.Freshwater Bryozoan | Home & Garden Information …

Url:https://hgic.clemson.edu/factsheet/freshwater-bryozoan/

1 hours ago Identification: Pectinatella magnifica is a species of freshwater bryozoan in the class Phylactolaemata. Like other species of bryozoans (also known as Ectoprocta or commonly as …

3.magnificent bryozoan (Pectinatella magnifica) - Species …

Url:https://nas.er.usgs.gov/queries/factsheet.aspx?SpeciesID=2335

17 hours ago Montz says bryozoans are quite common in many Minnesota waters, ranging from large rivers to lakes to small ponds. They are not toxic, venomous, or harmful. They don't really seem to …

4.Bryozoa - Wikipedia

Url:https://en.wikipedia.org/wiki/Bryozoa

35 hours ago Where are bryozoan colonies found? marine habitats. Bryozoans – also known as moss animals and sea mats – are adapted to live in many marine habitats, from the intertidal zone to the …

5.Bryozoans | GeoKansas - University of Kansas

Url:https://geokansas.ku.edu/bryozoans

19 hours ago

6.Bryozoa - Digital Atlas of Ancient Life

Url:https://www.digitalatlasofancientlife.org/learn/bryozoa/

12 hours ago

7.Gelatinous balls, moss animals and Bryozoans

Url:https://www.dnr.state.mn.us/areas/fisheries/baudette/bryozoans.html

5 hours ago

8.ADW: Bryozoa: INFORMATION - Animal Diversity Web

Url:https://animaldiversity.org/accounts/Bryozoa/

12 hours ago

A B C D E F G H I J K L M N O P Q R S T U V W X Y Z 1 2 3 4 5 6 7 8 9